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寒温带不同类型退化针叶林结构特征研究

Stand Structural Characteristics of Different Types of Degraded Coniferous Forests in Cold-Temperate Zone

  • 摘要:
    目的 阐明寒温带不同类型退化针叶林的非空间和空间结构特征,为该地区退化针叶林的结构和功能恢复提供理论依据。
    方法 在大兴安岭根河林区,选取退化近熟针叶林,以及不同类型火烧和采伐干扰的退化林分,共设置5块样地。调查样地内乔木、灌木和草本植物,分析各植被层物种多样性,运用林分空间结构分析方法分析林分空间结构参数(角尺度、混交度、密集度和大小比数)的一元、二元分布,计算林分结构多样性指数。
    结果 1)退化近熟针叶林样地林木较为稀疏,死亡树木株数占比较高;火烧干扰后的样地径级结构呈钟型,中小径级树木不足;采伐干扰退化针阔混交林样地中林木密度较高,兴安落叶松占比不足,低于40%。全部样地中,乔木层物种单一,仅有兴安落叶松和白桦两个树种;灌木多样性在受干扰较小的林分中较高,草本植物多样性则在受干扰较严重的林分中较高。2)随机木(Wi = 0.5)在全部样地中占据绝对优势地位,占比均超过55%,均匀木(Wi = 0或Wi = 0.25)占比最少。退化近熟针叶林样地、火烧干扰退化针叶林和采伐干扰退化针阔混交林样地平均角尺度分别为0.514,0.494和0.508,属于随机分布;火烧干扰重度退化林和火烧干扰退化针阔混交林样地平均角尺度分别达到0.563和0.540,属于聚集分布。5块样地混交度(Mi)均较低,以零度混交(Mi = 0)为主。所有样地兴安落叶松平均大小比数均小于0.5,兴安落叶松处于优势地位。所有样地中邻体随机分布且零度混交(Wi = 0.5且Mi = 0)的林木占比最大。3)采伐干扰退化针阔混交林样地的林分结构多样性达到2.033,火烧干扰重度退化林样地仅为0.248,林分结构多样性较低。
    结论 火烧干扰导致林分中小径级树木匮乏,天然更新受阻,径级结构不稳定;火烧和采伐干扰在一定程度上改变林分空间分布格局,降低林分结构多样性。因此对于火烧干扰林分,可采取人工措施破除更新障碍,促进天然更新;对于结构退化的林分,可应用结构化森林经营技术优化空间配置,提升结构多样性,从而推动寒温带退化针叶林结构修复与重建。

     

    Abstract:
    Objective This study aimed to elucidate the non-spatial and spatial structural characteristics of different types of degraded coniferous forests in cold-temperate zone, thereby providing a theoretical basis for the structural and functional restoration of degraded coniferous forests in this region.
    Method In the Genhe forest region of the Greater Khingan Mountains, 5 sample plots were established in degraded near-mature coniferous forests, as well as in stands subjected to fire and logging disturbance. Within each plot, trees, shrubs, and herbaceous plants were investigated to assess species diversity across different vegetation layers. Stand spatial structure analysis methods were employed to examine the univariate and bivariate distribution of the stand spatial structure parameters (uniform angle index, mingling, crowding, and neighborhood comparison) and structural diversity.
    Result 1) In the degraded near-mature coniferous forest plot, tree density was relatively low, with a high proportion of dead individuals. Stands affected by fire disturbance exhibited a unimodal (bell-shaped) diameter distribution, characterized by a deficiency of small- and medium-sized trees. In the Logging-degraded coniferous and broad-leaved mixed forest plot, the proportion of Larix gmelinii was less than 40of the tree layer, which consisted of Larix gmelinii and Betula platyphylla. Shrub diversity was higher in stands with less disturbance, while herbaceous plant diversity was greater in stands with more severe disturbance. 2) Randomly distributed trees (Wi = 0.5) overwhelmingly dominated all sample plots, accounting for more than 55% in each plot, while evenly distributed trees (Wi = 0 or Wi = 0.25) represented the samllest proportion. The mean uniform angle index for the degraded near-mature coniferous forest plot, the fire-degraded coniferous forest plot, and the logging-degraded coniferous and broad-leaved mixed forest plot were 0.514, 0.494, and 0.508, respectively, indicating a random distribution. While the fire severely degraded forest plot and the fire-degraded coniferous and broad-leaved mixed forest plot had mean uniform angle index values of 0.563 and 0.540, respectively, reflecting a cluster distribution. Mingling (Mi) was generally low across all plots, with zero-degree mingling (Mi = 0) being dominant. The average neighborhood comparison of Larix gmelinii in all plots was less than 0.5, indicating its dominant position. Trees with neighbors exhibiting both random distribution (Wi = 0.5) and zero-degree mingling (Mi = 0) accounted for the largest proportion in each plot. 3) Structural diversity varied markedly among forest types. The logging-degraded coniferous and broad-leaved mixed forest plot exhibited the highest structural diversity (2.033), whereas the fire severely degraded forest plot showed extremely low structural diversity (0.248).
    Conclusion Fire disturbance has led to a shortage of small-diameter trees within forest stands, hindered natural regeneration, and resulted in an unstable diameter distribution. Both fire and logging disturbances have altered the spatial distribution patterns of stands to some extent and reduced structural diversity. Therefore, for fire-disturbed stands, artificial interventions can be implemented to overcome regeneration barriers and facilitate natural regeneration. For structurally degraded stands, structured-based forest management techniques can be applied to optimize spatial structure, enhance structural diversity, and thereby promote the structural repair and reconstruction of degraded coniferous forests in cold-temperate zone.

     

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