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不同抚育采伐强度下竞争对华北落叶松人工林林木生长的调控

Regulation of Tree Growth by Competition under Different Thinning Intensities in Larix gmelinii var. principis-rupprechtii Plantations

  • 摘要:
    目的 阐明抚育采伐强度与邻域竞争对华北落叶松(Larix gmelinii var. principis-rupprechtii (Mayr) Pilg.)人工林保留木多维度生长的复合影响,揭示采伐对竞争效应的调节作用,以期为人工林的精准密度管理提供依据。
    方法 在山西省管涔山林区设立24块华北落叶松人工林固定监测样地(20 m × 20 m),于2017年末实施抚育采伐。设置两类采伐方案:(1)基于断面积比例,设置对照、15%强度采伐、30%强度采伐和45%强度采伐;(2)基于保留木密度,设置对照、保留1 800株·hm−2、保留1 350株·hm−2和保留900株·hm−2。基于2017年和2024年的两期每木检尺数据,量化单木胸径、树高、冠幅和地上生物量的年平均生长量,以及高径比和冠径比的变化量。构建线性模型或线性混合效应模型,解析抚育采伐强度、抚育采伐后的初始竞争及其交互作用对各项响应变量的影响,并将林木初始状态因子作为协变量加以控制。
    结果 (1)抚育采伐强度显著促进了保留木的胸径生长和生物量积累,且该效应表现出明显的个体初始状态依赖性。(2)邻域竞争极显著抑制了树高和冠幅生长,并诱导高径比和冠径比显著增加,导致林木形态稳定性下降及干形细长化。(3)随着抚育采伐强度增加,邻域竞争对树高和冠幅生长量的抑制作用呈现极显著增强趋势。
    结论 华北落叶松保留木的生长响应是采伐直接释放资源与间接改变竞争格局共同作用的结果。高强度采伐在显著促进径向生长的同时,可能因加剧保留木间的竞争而抑制高生长、影响树形。因此,在生产中需兼顾密度控制与空间配置,在采取较低保留密度的同时,应通过优化保留木的空间布局来缓解局部竞争压力,以实现大径材培育与林分稳定性维护的双重目标。

     

    Abstract:
    Objective This study aimed to clarify the combined effects of thinning intensity and neighborhood competition on the multi-dimensional growth of residual trees in Larix gmelinii var. principis-rupprechtii (Mayr) Pilg. plantations, and to reveal how thinning modulates competition effects, so as to provide a scientific basis for precise density management of planted forests.
    Methods Twenty-four permanent monitoring plots (20 m × 20 m) were established in Larix gmelinii var. principis-rupprechtii plantations in the Guancen Mountain Forest Region, Shanxi Province, China. Thinning treatments were implemented in late 2017 under two schemes: (1) Based on basal area ratio: control, 15% thinning intensity, 30% thinning intensity, and 45% thinning intensity; (2) Based on residual tree density: control, 1800 trees·hm−2, 1350 trees·hm−2, and 900 trees·hm−2. Using full-inventory measurements from 2017 and 2024, we quantified the annual mean increments of individual tree diameter at breast height (DBH), tree height, crown width, and aboveground biomass, as well as the changes in height-to-DBH ratio and crown-to-DBH ratio. Linear models and linear mixed effects models were constructed to analyze the effects of thinning intensity, initial post-thinning competition, and their interaction on each response variable, with initial tree characteristics included as covariates.
    Results (1) Thinning intensity significantly promoted DBH growth and biomass accumulation of residual trees, and this effect depended strongly on initial tree size. (2) Neighborhood competition highly significantly suppressed height and crown width growth, and significantly increased both the height-to-DBH ratio and crown-to-DBH ratio, indicating reduced morphological stability and slender stem form. (3) The inhibitory effect of neighborhood competition on height and crown width increments became highly significantly stronger as thinning intensity increased.
    Conclusion Growth responses of residual Larix gmelinii var. principis-rupprechtii trees reflect the combined outcomes of direct resource release due to thinning and indirect changes in competitive regimes. Although heavy thinning significantly promotes radial growth, it may intensify competition among residual trees, thereby limiting height growth and influencing tree form. Therefore, practical management should balance density control and spatial configuration, while adopting relatively low residual densities, managers should optimize the spatial distribution of retained trees to alleviate local competition, thereby supporting both large-diameter timber production and stand stability.

     

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